1. Colpo A., Molinari A., Boldrini P., Zivcak M., Brestic M., Demaria S., Baldisserotto C., Pancaldi S., Ferroni L. (2023) Thylakoid membrane appression in the giant chloroplast of Selaginella martensii Spring: a lycophyte challenges grana paradigms in shade-adapted species. Plant Science. Doi: 10.1016/j.plantsci.2023.111833.
  2. Colpo A., Demaria S., Boldrini P., Baldisserotto C., Pancaldi S., Ferroni L. (2023) Ultrastructural organization of the thylakoid system during the afternoon relocation of the giant chloroplast in Selaginella martensii Spring (Lycopodiophyta). Protoplasma. Doi: 10.1007/s00709-023-01888-w.
  3. Marrocchino E., Paletta M.G., Ferroni L., Manfrini R., Telloli C. (2023) Contribution of geosciences to cultural heritage conservation assessment: The case study of the Loggiato dei Cappuccini in Comacchio (Italy). Geosciences 13, 157. Doi: 10.3390/geosciences13060157.
  4. Baldisserotto C., Demaria S., Arcidiacono M., Benà E., Giacò P., Marchesini R., Ferroni L., Benetti L., Zanella M., Benini A., Pancaldi S. (2023) Enhancing urban wastewater treatment through isolated Chlorella strain-based phytoremediation in centrate stream: An analysis of algae morpho-physiology and nutrients removal efficiency. Plants 12, 1027. Doi: 10.3390/plants12051027.
  5. Colpo A., Demaria S., Baldisserotto C., Pancaldi S., Brestic M., Zivcak M.*, Ferroni L.* (2023) Long-Term Alleviation of the Functional Phenotype in Chlorophyll-Deficient Wheat and Impact on Productivity: A Semi-Field Phenotyping Experiment. Plants 18, 822. Doi: 10.3390/plants12040822. [*, corresponding authors]
  6. Ferroni L.*, Zivcak M., Kovar M., Colpo A., Pancaldi S., Allakhverdiev S.I., Brestic M.* (2022) Fast chlorophyll a fluorescence induction (OJIP) phenotyping of chlorophyll-deficient wheat suggests that an enlarged acceptor pool size of Photosystem I helps compensate for a deregulated photosynthetic electron flow. Journal of Photochemistry and Photobiology B - Biology 234, 112549. Doi:10.1016/j.jphotobiol.2022.112549. [*, corresponding authors]
  7. Baldisserotto C., Sabia A., Giovanardi M., Ferroni L., Maglie M., Pancaldi S. (2022) Chlorophyta microalgae as dietary protein supplement: a comparative analysis of productivity related to photosynthesis Chlorophyta microalgae as dietary protein supplement: a comparative analysis of productivity related to photosynthesis. Journal of Applied Phycology 34, 1323-1340. Doi:10.1007/s10811-022-02724-z.
  8. Filacek A., Zivcak M.*, Ferroni L.*, Barboricova M., Gasparovic K., Yang X., Landi M., Brestic M. (2022) Pre-acclimation to elevated temperature stabilizes the activity of photosystem I in wheat plants exposed to an episode of severe heat stress. Plants 11, 616. Doi:10.3390/plants11050616. [*, corresponding authors]
  9. Colpo A., Baldisserotto C., Pancaldi S., Sabia A., Ferroni L.* (2022) Photosystem II photoinhibition and photoprotection in a lycophyte, Selaginella martensii. Physiologia Plantarum 174, e13604. Doi: 10.1111/ppl.13604. [*, corresponding author]
  10. Maglie M., Baldisserotto C., Guerrini A. Sabia A., Ferroni L., Pancaldi S. (2021) A co-cultivation process of Nannochloropsis oculata and Tisochrysis lutea induces morpho-physiological and biochemical variations potentially useful for biotechnological purposes. Journal of Applied Phycology. Doi: 10.1007/s10811-021-02511-2.
  11. Ferroni L.*, Brestic M., Zivcak M., Cantelli R., Pancaldi S. (2021) Increased photosynthesis from a deep-shade to high-light regime occurs by enhanced CO2 diffusion into the leaf of Selaginella martensiiPlant Physiology and Biochemistry, 160, 143-154. Doi: 10.1016/j.plaphy.2021.01.012. [*, corresponding author]
  12. Baldisserotto C., Sabia A., Guerrini A., Demaria S., Maglie M., Ferroni L., Pancaldi S. (2021) Mixotrophic cultivation of Thalassiosira pseudonana with pure and crude glycerol: impact on lipid profileAlgal Research, 54, 102194. Doi: 10.1016/j.algal.2021.102194.
  13. Ferroni L.*, Colpo A., Baldisserotto C., Pancaldi S. (2021) In an ancient vascular plant the intermediate relaxing component of NPQ depends on a reduced stroma: Evidence from dithiothreitol treatment. Journal of Photochemistry and Photobiology – B Biology, 215, 112114. Doi: 10.1016/j.jphotobiol.2020.112114. [*, corresponding author]
  14. Baldisserotto C., Demaria S., Accoto O., Marchesini R., Zanella M., Benetti L., Avolio F., Maglie M., Ferroni L., Pancaldi S. (2020) Removal of nitrogen and phosphorus from thickening effluent of an urban wastewater treatment plant by an isolated green microalgaPlants, 9, 1802. doi: 10.3390/plants9121802.
  15. Ferroni L.*, Zivcak M.*, Sytar O.*, Kovar M., Watanabe N., Pancaldi S., Baldisserotto C., Brestic M. (2020) Chlorophyll-depleted wheat mutants are disturbed in photosynthetic electron flow regulation but can retain an acclimation ability to a fluctuating light regime. Environmental and Experimental Botany, 178, 104156. doi: 10.1016/j.envexpbot.2020.104156. [*, corresponding authors]
  16. Baldisserotto C., Sabia A., Ferroni L., Pancaldi S. (2019) Biological aspects and biotechnological potential of marine diatoms in relation to different light regimens. World Journal of Microbiology and Biotechnology, 35, 35. doi: 10.1007/s11274-019-2607-z.
  17. Ferroni L.*, Giovanardi M.*, Poggioli M., Baldisserotto C., Pancaldi S. (2018) Enhanced photosynthetic linear electron flow in mixotrophic green microalga Ettlia oleoabundans UTEX 1185. Plant Physiology and Biochemistry, 130, 215-223. doi: 10.1016/j.plaphy.2018.07.005 [*, equal contribution]
  18. Giovanardi M.*, Pantaleoni L.*, Ferroni L.*, Pagliano C., Albanese P., Baldisserotto C., Pancaldi S. (2018) In pea stipules a functional photosynthetic electron flow occurs despite a reduced dynamicity of LHCII association with photosystems. Biochimica et Biophysica Acta - Bioenergetics, 1859, 1025-1038. doi: 10.1016/j.bbabio.2018.05.013 [*, equal contribution]
  19. Ferroni L.*, Cucuzza S., Angeleri M., Aro E-M., Pagliano C., Giovanardi M., Baldisserotto C., Pancaldi S. (2018) In the lycophyte Selaginella martensii is the "extra-qT" related to energy spillover? Insights into photoprotection in ancestral vascular plants. Environmental and Experimental Botany, 154, 110-122. doi: [*, corresponding author]
  20. Giovanardi M., Poggioli M., Ferroni L., Lespinasse M., Aro E-M., Baldisserotto C., Pancaldi S. (2017) Higher packing of thylakoid complexes ensures a preserved Photosystem II activity in mixotrophic Neochloris oleoabundans. Algal Research, 25, 322-332. doi: 10.1016/j.algal.2017.05.020.
  21. Kalaji H.M., Schansker G., Brestic M., Bussotti F., Calatayud A., Ferroni L., Goltsev V., Guidi L., Jajoo A., Li P., Losciale P., Mishra V.K., Misra A.N., Nebauer S.G., Pancaldi S., Penella C., Pollastrini M., Suresh K., Tambussi E., Yanniccari M., Zivcak M., Cetner M.G., Samborska I.A., Stirbet A., Olsovska K., Kunderlikova K., Shelonzek H., Rusinowski A., Baba W. (2017) Frequently asked questions about chlorophyll fluorescence, the sequel. Photosynthesis Research, 132, 13-66. doi: 10.1007/s11120-016-0318-y
  22. Ferroni L.*, Suorsa M., Aro E-M., Baldisserotto C., & Pancaldi S. (2016) Light acclimation in the lycophyte Selaginella martensii depends on changes in the amount of photosystems and on the flexibility of light-harvesting complex II antenna association to both photosystems. New Phytologist, 211, 554–568. doi: 10.1111/nph.13939. [*, corresponding author]
  23. Baldisserotto C., Popovich C., Giovanardi M., Sabia A., Ferroni L., Constenla D., Leonardi P.I., & Pancaldi S. (2016) Photosynthetic aspects and lipid profiles in the mixotrophic alga Neochloris oleoabundans as useful parameters for biodiesel production. Algal Research, 16, 255-265. doi: 10.1016/j.algal.2016.03.022
  24. Giovanardi M., Baldisserotto C., Daglia M., Ferroni L., Sabia A., & Pancaldi S. (2016) Morpho-physiological aspects ofScenedesmus acutus PVUW12 cultivated with a dairy industry waste and after starvation. Plant Biosystems, 150, 767-775. doi: 10.1080/11263504.2014.991361
  25. Reale L., Ferranti F., Mantilacci S., Corboli M., Aversa S., Landucci F., Baldisserotto C., Ferroni L., Pancaldi S., & Venanazoni R. (2016) Cyto-histological and morpho-physiological responses of common duckweed (Lemna minor L.) to chromium. Chemosphere 145, 98-105. doi: 10.1016/j.chemosphere.2015.11.047
  26. Sabia A., Baldisserotto C., Biondi S., Marchesini R., Tedeschi P., Maietti A., Giovanardi M., Ferroni L., & Pancaldi S. (2015) Re-cultivation of Neochloris oleoabundans in exhausted autotrophic and mixotrophic media: the potential role of polyamines and free fatty acids. Applied Microbiology and Biotechnology 99, 10597-10609. doi: 10.1007/s00253-015-6908-3
  27. Ferroni L.*, Angeleri M., Pantaleoni L., Pagliano C., Longoni P., Marsano F., Aro E-M., Suorsa M., Baldisserotto C., Giovanardi M., Cella R., & Pancaldi S. (2014) Light-dependent reversible phosphorylation of the minor photosystem II antenna Lhcb6 (CP24) occurs in lycophytes. The Plant Journal 77, 893-905. doi: 10.1111/tpj.12437. [*, corresponding author]
  28. Kalaji H.M., Schansker G., Ladle R.J., Goltsev V., Bosa K., Allakhverdiev S.I., Brestic M., Bussotti F., Calatayud A., Dąbrowski P., Elsheery N.I., Ferroni L., Guidi L., Hogewoning S. W., Jajoo A., Misra A.N., Nebauer S.G., Pancaldi S., Penella C., Poli DB., Pollastrini M., Romanowska-Duda Z.B., Rutkowska B., Serôdio J., Suresh K., Szulc W., Tambussi E., Yanniccari M., Zivcak M. (2014) Frequently asked questions about in vivo chlorophyll fluorescence: practical issues. Photosynthesis Research 121, 122-158. doi: 10.1007/s11120-014-0024-6
  29. Pantaleoni L., Longoni L., Ferroni L., Baldisserotto C., Leelavathi S., Reddy V.S., Pancaldi S., & Cella R. (2014) Chloroplast molecular farming: efficient production of a thermostable xylanase by Nicotiana tabacum plants and long-term conservation of the recombinant enzyme. Protoplasma 251, 639-648. doi: 10.1007/s00709-013-0564-1.
  30. Baldisserotto C., Giovanardi M., Ferroni L., & Pancaldi S. (2014) Growth, morphology and photosynthetic responses of Neochloris oleoabundans during cultivation in a mixotrophic brackish medium and subsequent starvation. Acta Physiologiae Plantarum 36, 461-472. doi: 10.1007/s11738-013-1426-3.
  31. Giovanardi M., Baldisserotto C., Ferroni L., Longoni P., Cella R., & Pancaldi S. (2014) Growth and lipid synthesis promotion in mixotrophic Neochloris oleoabundans (Chlorophyta) cultivated with glucose. Protoplasma 251, 115-125 doi: 10.1007/s00709-013-0531-x.
  32. Baldisserotto C.*, Ferroni L.*, Pantaleoni L., & Pancaldi S. (2013) Comparison of photosynthesis recovery dynamics in floating leaves of Trapa natans after inhibition by manganese or molybdenum: Effects on Photosystem II. Plant Physiology and Biochemistry 70, 387-395. doi: 10.1016/j.plaphy.2013.05.044. [*, equal contribution]
  33. Giovanardi M., Ferroni L., Baldisserotto C., Tedeschi P., Maietti A., Pantaleoni L., & Pancaldi S. (2013) Morphophysiological analyses of Neochloris oleoabundans (Chlorophyta) grown mixotrophycally in a carbon-rich waste product. Protoplasma 250, 161-174.
  34. Ferroni L.*, Pantaleoni L.*, Baldisserotto C., Aro E.-M., & Pancaldi S. (2013) Low photosynthetic activity is linked to changes in the organization of photosystem II in the fruit of Arum italicum. Plant Physiology and Biochemistry 63, 140-150. [*, equal contribution]
  35. Baldisserotto C., Ferroni L., Giovanardi M., Boccaletti L., Pantaleoni L., & Pancaldi S. (2012) Salinity promotes growth of freshwater Neochloris oleoabundans UTEX 1185 (Sphaeropleales, Chlorophyta): morphophysiological aspects. Phycologia 51 (6), 700-710.
  36. Ferroni L., Baldisserotto C., Giovanardi M., Pantaleoni L., Morosinotto T., & Pancaldi S. (2011) Revised assignment of room-temperature chlorophyll fluorescence emission bands in single living cells of Chlamydomonas reinhardtii. Journal of Bioenergetics and Biomembranes 43, 163-173.
  37. Baldisserotto C., Ferroni L., Zanzi C., Marchesini R., Pagnoni A., & Pancaldi S. (2010) Morpho-physiological and biochemical responses in the floating lamina of Trapa natans exposed to molybdenum. Protoplasma 240, 83-97.
  38. Ferroni L., Klisch M., Pancaldi S., & Häder D.-P. (2010) Complementary UV-absorption of mycosporine-like aminoacids and scytonemin is responsible for the UV-insensitivity of photosynthesis in Nostoc flagelliforme. Marine Drugs 8, 106-121.
  39. Pantaleoni L., Ferroni L., Baldisserotto C., Aro E-M., & Pancaldi S. (2009) Photosystem II organisation in chloroplasts of Arum italicum leaf depends on tissue location. Planta 230, 1019-1031.
  40. Ferroni L., Baldisserotto C., Fasulo M.P., Moro I., Cella R., & Pancaldi S. (2009) Changes in proplastid organization promoted by an inhibitor of the DNA-methyltransferase in dark-grown dividing Euglena gracilis. Plant Biosystems 143, 241-251.
  41. Ferroni L., Baldisserotto C., Pantaleoni L., Fasulo M.P., Fagioli P., & Pancaldi S. (2009) Degreening of the unicellular alga Euglena gracilis: thylakoid composition, room temperature fluorescence spectra and chloroplast morphology. Plant Biology 11, 631-641.
  42. Ferroni L., Baldisserotto C., Pantaleoni L., Billi P., Fasulo M.P., & Pancaldi S. (2007) High salinity alters chloroplast morpho-physiology in a freshwater Kirchneriella species (Selenastraceae) from Ethiopian Lake Awasa. American Journal of Botany 94, 1972-1983.
  43. Baldisserotto C., Ferroni L., Anfuso E., Pagnoni A., Fasulo M.P., & Pancaldi S. (2007) Responses of Trapa natans L. floating laminae to high concentrations of manganese. Protoplasma 231, 65-82.
  44. Ferroni L., Baldisserotto C., Zennaro V., Soldani C. Fasulo M.P. & Pancaldi S. (2007) Acclimation to darkness in the marine chlorophyte Koliella antarctica cultured under low salinity: hypotheses on its origin in the polar environment. European Journal of Phycology 42, 91-104.
  45. Baldisserotto C., Ferroni L., Moro I., Fasulo M.P. & Pancaldi S. (2005) Modulations of the thylakoid system in snow xanthophycean alga cultured in the dark for two months: comparison between microspectrofluorimetric responses and morphological aspects. Protoplasma 226, 125-135.
  46. Baldisserotto C., Ferroni L., Andreoli C., Fasulo M.P., Bonora A. & Pancaldi S. (2005) Dark-acclimation of the chloroplast in Koliella antarctica exposed to a simulated austral night condition. Arctic, Antarctic and Alpine Research 37, 146-156.
  47. Ferroni L., Baldisserotto C., Fasulo M.P., Pagnoni A. & Pancaldi S. (2004) Adaptive modifications of the photosynthetic apparatus in Euglena gracilis Klebs exposed to manganese excess. Protoplasma 224, 167-177.
  48. Baldisserotto C., Ferroni L., Medici V., Pagnoni A., Pellizzari M., Fasulo M.P., Fagioli F., Bonora A. & Pancaldi S. (2004) Specific intra-tissue responses to manganese in the floating lamina of Trapa natans L.. Plant Biology 6, 578-589.
  49. Pancaldi S., Baldisserotto C., Ferroni L., Bonora A. & Fasulo M.P. (2002) Room temperature microspectrofluorimetry as a useful tool for studying the assembly of the PSII chlorophyll-protein complexes in single living cells of etiolated Euglena gracilis Klebs during the greening process. Journal of Experimental Botany 53, 1753-1763.



  1. Baldisserotto C., Ferroni L., Felisati B., Fasulo M.P., Pancaldi S. (2005) Studio microspettrofluorimetrico ed ultrastrutturale delle forme vegetative di un ceppo nivale di Chlorella saccharophila. In: Le Scienze Botaniche nel XXI secolo. 100° Congresso della Società Botanica Italiana. INFORMATORE BOTANICO ITALIANO 37/1 B, 622-623.
  2. Ferroni L. (2012) Photosynthetic Acclimation to the Light Environment: Molecular Mechanisms to Understand Plant Consortia. JOURNAL OF ECOSYSTEM AND ECOGRAPHY, 2:e104. doi:10.4172/2157-7625.1000e104. [invited editorial]



  1. Pancaldi S., Baldisserotto C., Ferroni L., Pantaleoni L. (2023) Fondamenti di Botanica Generale - teoria e pratica in laboratorio. Terza Edizione. MILANO: McGraw-Hill Education, ISBN: 9788838695360.
  1. Pancaldi S., Baldisserotto C., Ferroni L., Pantaleoni L. (2019) Fondamenti di Botanica Generale - teoria e pratica in laboratorio. Seconda Edizione. MILANO: McGraw-Hill Education, ISBN: 9788838695360.
  2. Pancaldi S., Baldisserotto C., Ferroni L., Pantaleoni L. (2011) Fondamenti di Botanica Generale. Teoria e pratica. MILANO: McGraw-Hill, ISBN: 9788838665707.



  • Pancaldi S., Ferroni L., Baldisserotto C., Giovanardi M., Mai L. (2012) Impianto e procedimento per lo smaltimento del carico di azoto di reflui organici in allevamenti zootecnici. Brevetto d’invenzione industriale, concessione n. 001412445 (domanda del 15.12.2012, concessione del 27.11.2014). Titolarità UniFE.